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    Resource availability and plant antiherbivore defense. Science : — Influence of female cones on the vegetative growth of Pinus contorta trees.

    Tree Physiology 6 : — R package version 1. Google Preview. Relation between cone production and diameter increment of Douglas fir Pseudotsuga memiesii Mirb.

    Franco , grand fir Abies grandis Dougl. Canadian Journal of Botany 43 : — ArcGIS desktop: release Influence of region of provenance and climate factors on wood anatomical traits of Pinus nigra Arn.

    European Journal of Forest Research : — To grow or defend? Pine seedlings grow less but induce more defences when a key resource is limited.

    Tree Physiology 35 : — Resin duct characteristics associated with tree resistance to bark beetles across lodgepole and limber pines.

    Ecological Applications 19 : — Forcella F Estimating pinyon cone production in New Mexico and western Oklahoma. The Journal of Wildlife Management 45 : — Fox J , Weisberg S.

    Thousand Oaks, CA : Sage. Anatomical and chemical defenses of conifer bark against bark beetles and other pests.

    The New Phytologist : — Fritts HC. Tree rings and climate , 1st edn. Cambridge, MA : Academic Press. Google Earth Engine: planetary-scale geospatial analysis for everyone.

    Remote Sensing of Environment : 18 — Gross HL. Crown deterioration and reduced growth associated with excessive seed production by birch. Canadian Journal of Botany 50 : — Costs of defense and their effects on plant productivity.

    In: Givnish TJ , ed. On the economy of plant form and function. Gurevitch, J. The carbon—nutrient balance hypothesis: its rise and fall.

    Ecology Letters 4 : 86 — Genetic parameters of growth and wood quality traits in Picea abies. Scandinavian Journal of Forest Research 19 : 14 — The dilemma of plants: to grow or defend.

    The Quarterly Review of Biology 67 : — Hood S , Sala A. Ponderosa pine resin defenses and growth: metrics matter.

    Importance of resin ducts in reducing ponderosa pine mortality from bark beetle attack. Mechanisms to mitigate the trade-off between growth and defense.

    The Plant Cell 29 : — Keeling CI , Bohlmann J. Genes, enzymes and chemicals of terpenoid diversity in the constitutive and induced defence of conifers against insects and pathogens.

    Resistance of Sitka spruce Picea sitchensis Bong. Forestry 84 : 83 — Kläy M. Are defensive structures good predictors of tree mortality under drought and insect pressure?

    Klutsch JG , Erbilgin N. Dwarf mistletoe infection in jack pine alters growth-defense relationships. Tree Physiology 38 : — Scale of mast-seeding and tree-ring growth.

    Nature : — Patterns of annual seed production by Northern hemisphere trees: a global perspective. The American Naturalist : 59 — Koricheva J. Ecology 83 : — Genetic variation in space and time in a population of ponderosa pine.

    Heredity 46 : — Environmental effects on constitutive and inducible resin defences of Pinus taeda. Ecology Letters 3 : — Loomis WE.

    Growth-differentiation balance vs carbohydrate-nitrogen ratio. Proceedings of the American Society for Horticultural Science 29 : — Lorio PL. Growth-differentiation balance: a basis for understanding southern pin beetle-tree interactions.

    Forest Ecology and Management 14 : — Variation in wood anatomical traits of Pinus sylvestris L. Trees Structure and Function 24 : — The New Phytologist : 91 — Monks A , Kelly D.

    Austral Ecology 31 : — Masting in ponderosa pine: comparisons of pollen and seed over space and time. Differential allocation of constitutive and induced chemical defenses in pine tree juveniles: a test of the optimal defense theory.

    PLoS One 7 : e Differentiation of persistent anatomical defensive structures is costly and determined by nutrient availability and genetic growth-defence constraints.

    Moustakas A , Evans MR. Effects of growth rate, size, and light availability on tree survival across life stages: a demographic analysis accounting for missing values and small sample sizes.

    BMC Ecology 15 : 6. Neale DB , Kremer A. Forest tree genomics: growing resources and applications. Nature Reviews Genetics 12 : — Niinemets Ü.

    A review of light interception in plant stands from leaf to canopy in different plant functional types and in species with varying shade tolerance.

    Ecological Research 25 : — Obeso JR. The costs of reproduction in plants. Owens JN. The relative importance of initiation and early development on cone production in Douglas-fir.

    Canadian Journal of Botany 47 : — Ecosphere 9 : e Mechanisms of mast seeding: resources, weather, cues, and selection.

    R package version 3. Oregon State University. R Core Team. R: a language and environment for statistical computing. Version 3. Declines in pinyon pine cone production associated with regional warming.

    Ecosphere 3 : Ecosphere 8. Woodland resilience to regional drought: dominant controls on tree regeneration following overstorey mortality. Journal of Ecology : — A robust method to determine historical annual cone production among slow-growing conifers.

    Forest Ecology and Management : 1 — 6. Resin yield in Pinus pinaster is related to tree dendrometry, stand density and tapping-induced systemic changes in xylem anatomy.

    Forest Ecology and Management : 47 — Ronco FP Jr. Pinus edulis Engelm. Silvics of North America, Volume 1. Rosner S , Hannrup B. Resin canal traits relevant for constitutive resistance of Norway spruce against bark beetles: environmental and genetic variability.

    Forest Ecology and Management : 77 — Masting in whitebark pine Pinus albicaulis depletes stored nutrients.

    Costs of constitutive and herbivore-induced chemical defences in pine trees emerge only under low nutrient availability. Journal of Ecology 99 : — Stamp N.

    Out of the quagmire of plant defense hypotheses. The Quarterly Review of Biology 78 : 23 — Introduction to tree-ring dating , 2nd edn.

    Tappeiner JC. Effect of cone production on branch, needle, and xylem ring growth of Sierra Nevada Douglas-fir. Forest Science 15 : — Trapp S , Croteau R.

    Defensive resin biosynthesis in conifers. Trade-offs between growth, reproduction and defense in response to resource availability manipulations.

    PLoS One 13 : e Resource allocation on dwarf shoots of birch Bentla pendula : reproduction and leaf growth. The New Phytologist 91 : — Discovering candidate genes that regulate resin canal number in Pinus taeda stems by integrating genetic analysis across environments, ages, and populations.

    Temperature as a potent driver of regional forest drought stress and tree mortality. Nature Climate Change 3 : — In prep.

    An year record of decadal-scale hydroclimatic variability in the Upper Arkansas River basin from bristlecone pine.

    Quaternary Research. Intraspecific variation of anatomical and chemical defensive traits in maritime pine Pinus pinaster as factors in susceptibility to the pinewood nematode Bursaphelenchus xylophilus.

    Trees 29 : — Züst T , Agrawal AA. Trade-offs between plant growth and defense against insect herbivory: an emerging mechanistic synthesis.

    Annual Review of Plant Biology 68 : — Mixed effects models and extensions in ecology with R. New York: Springer. Oxford University Press is a department of the University of Oxford.

    It furthers the University's objective of excellence in research, scholarship, and education by publishing worldwide.

    Sign In or Create an Account. Sign In. Advanced Search. Search Menu. Skip Nav Destination Article Navigation. Close mobile search navigation Article Navigation.

    Volume Article Contents Abstract. Materials and Methods. Supporting Information. Sources of Funding. Conflict of Interest.

    Literature Cited. Article Navigation. Redmond colostate. Oxford Academic. Thomas Seth Davis. Scott Ferrenberg. Andreas P Wion. Select Format Select format.

    Permissions Icon Permissions. Abstract The cost of plant reproduction or defence at the expense of other fitness traits is a central component of life history theory.

    Defence; masting , mast seeding , Pinus edulis , pinyon pine , reproduction , resource allocation , trade-offs , tree growth.

    Open in new tab Download slide. Table 1. This metric was standardized by the length of the annual ring sampled, which varied slightly across tree cores, and thus total resin duct area is: resin duct area mm per year per sample length mm.

    Open in new tab. Google Scholar Crossref. Search ADS. Google Scholar PubMed. Issue Section:. Download all slides. Supplementary data. Comments 0.

    Add comment Close comment form modal. I agree to the terms and conditions. You must accept the terms and conditions.

    Add comment Cancel. Submit a comment. The MFD can also be used to view other non-overlay type of data e. EICAS displays are often designed to mimic traditional round gauges while also supplying digital readouts of the parameters.

    EICAS improves situational awareness by allowing the aircrew to view complex information in a graphical format and also by alerting the crew to unusual or hazardous situations.

    For example, if an engine begins to lose oil pressure, the EICAS might sound an alert, switch the display to the page with the oil system information and outline the low oil pressure data with a red box.

    Unlike traditional round gauges, many levels of warnings and alarms can be set. Proper care must be taken when designing EICAS to ensure that the aircrew are always provided with the most important information and not overloaded with warnings or alarms.

    EFIS provides pilots with controls that select display range and mode for example, map or compass rose and enter data such as selected heading.

    Where other equipment uses pilot inputs, data buses broadcast the pilot's selections so that the pilot need only enter the selection once.

    For example, the pilot selects the desired level-off altitude on a control unit. The EFIS repeats this selected altitude on the PFD, and by comparing it with the actual altitude from the air data computer generates an altitude error display.

    This same altitude selection is used by the automatic flight control system to level off, and by the altitude alerting system to provide appropriate warnings.

    The EFIS visual display is produced by the symbol generator. This receives data inputs from the pilot, signals from sensors, and EFIS format selections made by the pilot.

    The symbol generator can go by other names, such as display processing computer, display electronics unit, etc. The symbol generator does more than generate symbols.

    It has at the least monitoring facilities, a graphics generator and a display driver. The required computations are performed, and the graphics generator and display driver produce the inputs to the display units.

    Like personal computers, flight instrument systems need power-on-self-test facilities and continuous self-monitoring.

    Flight instrument systems, however, need additional monitoring capabilities:. Traditional electromechanical displays are equipped with synchro mechanisms that transmit the pitch, roll, and heading shown on the captain and first officer's instruments to an instrument comparator.

    The comparator warns of excessive differences between the Captain and First Officer displays. Even a fault as far downstream [3] as a jam in, say, the roll mechanism of an ADI triggers a comparator warning.

    The instrument comparator thus provides both comparator monitoring and display monitoring. With EFIS, the comparator function is simple: Is roll data bank angle from sensor 1 the same as roll data from sensor 2?

    Comparison monitors give warnings for airspeed, pitch, roll, and altitude indications. More advanced EFIS systems have more comparator monitors.

    In this technique, each symbol generator contains two display monitoring channels. One channel, the internal, samples the output from its own symbol generator to the display unit and computes, for example, what roll attitude should produce that indication.

    Any difference has probably been introduced by faulty processing, and triggers a warning on the relevant display.

    The external monitoring channel carries out the same check on the symbol generator on the other side of the flight deck: the Captain's symbol generator checks the First Officer's, the First Officer's checks the Captain's.

    Whichever symbol generator detects a fault, puts up a warning on its own display. The external monitoring channel also checks sensor inputs to the symbol generator for reasonableness.

    A spurious input, such as a radio height greater than the radio altimeter's maximum, results in a warning. At various stages of a flight, a pilot needs different combinations of data.

    Ideally, the avionics only show the data in use—but an electromechanical instrument must be in view all the time.

    Under normal conditions, an EFIS might not display some indications, e. Only when some parameter exceeds its limits does the system display the reading.

    In the case of an input failure, an electromechanical instrument adds yet another indicator—typically, a bar drops across the erroneous data.

    EFIS, on the other hand, removes invalid data from the display and substitutes an appropriate warning.

    A de-clutter mode activates automatically when circumstances require the pilot's attention for a specific item. For example, if the aircraft pitches up or down beyond a specified limit—usually 30 to 60 degrees—the attitude indicator de-clutters other items from sight until the pilot brings the pitch to an acceptable level.

    This helps the pilot focus on the most important tasks. Traditional instruments have long used color, but lack the ability to change a color to indicate some change in condition.

    The electronic display technology of EFIS has no such restriction and uses color widely. For example, as an aircraft approaches the glide slope, a blue caption can indicate glide slope is armed, and capture might change the color to green.

    Typical EFIS systems color code the navigation needles to reflect the type of navigation. Magenta needles indicate GPS navigation.

    EFIS provides versatility by avoiding some physical limitations of traditional instruments.

    King Cone Eis They King Cone Eis also lighter, and Stolichnaya Elit a lower volume. In addition, we were unable to sample trees that had highly damaged branches and thus no available markers of cone production, which may have been exceptionally poor or high producers of resin ducts or cone production. Only when some parameter exceeds its limits does Naruto Spielen system display the reading. Under normal conditions, Lotto Systeme EFIS might not display some indications, e. This receives data inputs from the pilot, signals from sensors, and EFIS format selections Empire Online Spiel by the pilot. Crown deterioration and reduced growth associated with excessive seed production by birch. Spielbank Bad Füssing alerts Article activity alert. Effect of cone production on branch, needle, and xylem ring growth of Sierra Nevada Douglas-fir. Our The Dark King uses a unique data set of within-individual time series to provide new insights into allocations between physiological expressions of growth, defence and reproduction in a long-lived masting conifer species, and indicate trade-offs do occur between tree defence and reproduction among individuals but not between growth and defence. Subject alert. Yet few authors have considered the role of allocation to defence in understanding trade-offs Dfb Pokalspiel Heute mast-seeding species, likely because of the challenges of simultaneously measuring growth, defence and reproduction over time. Our seed cone production model assessed how two metrics of growth shoot and radial growth and two metrics of defence total resin duct area and Mahjong Quest resin duct size were associated with seed cone production see Table 1 for a description of all variables. The dilemma of plants: to grow or defend.
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